Publications
Original Research Articles
Pre-Print Martínez de Pinillos González M, Álvarez Fernández A, Delgado Esteban B, Delgado M, Dern LL, Irish JD, Kondo O, Martinón Torres M, Modesto-Mata M, Scott GR, Thiebaut A, Paul KS, Pilloud MA, Rathmann H, Reyes-Centeno H, Vlemincq-Mendieta T, Hlusko LJ. DeMoDa: A global open database for comparative research in human dental morphological variation. bioRxiv doi:10.64898/2026.05.30.728952
In Press Bosman AM, Buck LT, Reyes-Centeno H, Mirazón Lahr M, Stringer C, Grün R, Shao Q, and Harvati K. The morphological affinities of the fossil cranium from Kabua, Kenya. Journal of the Royal Anthropological Institute. doi:10.1111/1467-9655.70137 [Impact factor: 1.1]
2026 López Sosa MC, Reyes-Centeno H, Menéndez LP, Oliveira RE, Campi de Castro C, and Strauss A. Endocranial morphological affinities of the Early Holocene individuals from Lagoa Santa and implications for the peopling of the Americas. Boletim do Museu Paraense Emílio Goeldi: Ciências Humanas. 21(1):e20240059. doi:10.1590/2178-2547-BGOELDI-2024-0059
2025 Szeniczey T, Melis E, Jakab J, Bátora J, Major I, Molnár M, Horváth A, Palcsu L, Gémes A, Gyenesei K, Reyes-Centeno H, Kiss V, and Hajdu T. The population history of Central Europe in the Early Bronze Age (2200/2100–1600/1500 BC) - A craniometric approach. Archaeological and Anthropological Sciences 17, 217 (2025). doi:10.1007/s12520-025-02324-1
2025 Keppeler J, Rodriguez M, Koontz S, Wise A, Mink P, Crothers G, Murphy P, Robertson J, Reyes-Centeno H, and Uhl A. Remote sensing of American Revolutionary War fortification at Butts Hill (Portsmouth, Rhode Island). Heritage 8(10), 430 doi:10.3390/heritage8100430
2025 Adel M, Hunt KJ, Lau D, Hartsfield JK, Reyes-Centeno H, Beeman CS, Elshebiny T, and Sharab L. Precision assessment of facial asymmetry using 3D imaging and artificial intelligence. Journal of Clinical Medicine 14(20), 7172. doi:10.3390/jcm14207172
2023 Napora K, Crothers G, Hadden C, Guerre L, Waldman L, Reyes-Centeno H, Keppeler J, Imler M, Jakaitis E, Metz A, and Mink P. Multi-method analysis of an historic wooden trough from Kentucky, USA: A case study in supporting artifact oral histories with heritage science. Heritage Science 11(235). doi:10.1186/s40494-023-01075-3
2023 Rathmann H, Perretti S, Porcu V, Hanihara T, Scott RG, Irish J, Reyes-Centeno H, Ghirotto S, and Harvati K. Inferring human neutral genetic variation from craniodental phenotypes. PNAS Nexus 2(7). doi:10.1093/pnasnexus/pgad217
2022 Beaudet A, d’Errico F, Backwell L, Wadley L, Zipfel B, de la Peña P, Reyes-Centeno H. A reappraisal of the Border Cave 1 cranium (KwZulu-Natal, South Africa). Quaternary Science Reviews 282:107452. doi:10.1016/j.quascirev.2022.107452
2021 Arner AM, Grogan KE, Grabowski M, Reyes-Centeno H, and Perry G. Patterns of recent natural selection on genetic loci associated with sexually differentiated human body size and shape phenotypes. PLOS Genetics 17(6):e1009562. doi:10.1371/journal.pgen.1009562
2021 Karakostis FA, Reyes-Centeno H, Franken M, Hotz G, Rademaker K, and Harvati K. Bio-cultural evidence of precise manual activities in an Early Holocene individual of the high-altitude Peruvian Andes. American Journal of Physical Anthropology. 174(1):35-48. doi:10.1002/ajpa.24160
2020 Mori T, Profico A, Reyes-Centeno H, and Harvati K. Frontal bone virtual reconstruction and geometric morphometric analysis of the mid-Pleistocene hominin KNM-OG 45500 (Olorgesailie, Kenya). Journal of Anthropological Sciences 98:49-72. doi:10.4436/jass.98022
2020 Bosman A, Reyes-Centeno H, and Harvati K. A virtual assessment of the suprainiac depressions on the Eyasi I and ADU-VP-1/3 crania. Journal of Human Evolution 145:102815. doi:10.1016/j.jhevol.2020.102815
2020 Rathmann H and Reyes-Centeno H. Testing the utility of dental morphological trait combinations for inferring human neutral genetic variation. Proceedings of the National Academy of Sciences USA 117(20):10769-10777. doi:10.1073/pnas.1914330117
2019 Urban M, Reyes-Centeno H, Bellamy KR, and Pache MJ. The areal typology of western Middle and South America: Towards a comprehensive view. Linguistics 57(6):1403-1463 doi:10.1515/ling-2019-0032
2018 Posth C, Nakatsuka N, Lazaridis I, Skoglund P, Mallick S, Lamnidis TC, Rohland N, Nägele K, Adamski N, Bertolini E, Broomandkhoshbacht N, Cooper A, Culleton BJ, Ferraz T, Ferry M, Furtwängler A, Haak W, Harkins K, Harper TK, Hünemeier T, Lawson AM, Llamas B, Michel M, Nelson E, Oppenheimer J, Patterson N, Schiffels S, Sedig J, Stewardson K, Talamo S, Wang C-C, Hublin J-J, Hubbe M, Harvati K, Nuevo Delaunay A, Beier J, Francken M, Kaulicke P, Reyes-Centeno H, Rademaker K, Trask WR, Robinson M, Gutierrez SM, Prufer KM, Salazar-García DC, Chim EN, Müller Plumm Gomes L, Alves ML, Liryo A, Inglez M, Oliveira RE, Bernardo DV, Barioni A, Wesolowski V, Scheifler NA, Rivera MA, Plens CR, Messineo PG, Figuti L, Corach D, Scabuzzo C, Eggers S, DeBlasis P, Reindel M, Méndez C, Politis G, Tomasto-Cagigao E, Kennett DJ, Strauss A, Fehren-Schmitz L, Krause J, and Reich D. Reconstructing the Deep Population History of Central and South America. Cell 175(5):1185-1197. doi:10.1016/j.cell.2018.10.027
2017 Rathmann H, Reyes-Centeno H, Ghirotto S, Creanza N, Hanihara T, and Harvati K. Reconstructing human population history from dental phenotypes. Scientific Reports 7(1):12495. doi:10.1038/s41598-017-12621-y
2017 Reyes-Centeno H, Rathmann H, Hanihara T, and Harvati K. Testing modern human out-of-Africa dispersal models using dental non-metric data. Current Anthropology 58(S17):S406-417. doi:10.1086/694423
2017 Reyes-Centeno H, Ghirotto S., and Harvati K. Genomic validation of the differential preservation of population history in modern human cranial anatomy. American Journal of Physical Anthropology 162(1):170-179. doi:10.1002/ajpa.23060
2016 Reyes-Centeno H, Harvati K, Jäger G. Tracking modern human population history from linguistic and cranial phenotype. Scientific Reports 6:36645. doi:10.1038/srep36645
2016 Uhl A, Reyes-Centeno H, Grigorescu D, Kranioti EF, and Harvati K. Inner ear morphology of the Cioclovina early modern European calvaria from Romania. American Journal of Physical Anthropology 160(1):62-70. doi:10.1002/ajpa.22938
2015 Reyes-Centeno H, Hubbe M, Hanihara T, Stringer C, Harvati K. Testing modern human out-of-Africa dispersal models and implications for modern human origins. Journal of Human Evolution 87, 95-106. doi:10.1016/j.jhevol.2015.06.008
2014 Reyes-Centeno H, Ghirotto S, Détroit F, Grimaud-Hervé D, Barbujani G, and Harvati K. Genomic and cranial phenotype data support multiple modern human dispersals from Africa and a southern route into Asia. Proceedings of the National Academy of Sciences USA 111(20):7248-7253. doi:10.1073/pnas.1323666111
Reviews
2022 Harvati K and Reyes-Centeno H. Evolution of Homo in the Middle and Late Pleistocene. Journal of Human Evolution 173:103279. doi:10.1016/j.jhevol.2022.103279
2016 Reyes-Centeno H. Out of Africa and into Asia: Fossil and genetic evidence on modern human origins and dispersals. Quaternary International 416:249-262. doi:10.1016/j.quaint.2015.11.063
Book Chapters
2021 Evteev A, Santos P, Groshva A, Reyes-Centeno H, and Ghirotto S. Associations between human genetic and craniometric differentiation across North Eurasia: the role of geographic scale. In: Reyes-Centeno H and Harvati K (Eds.), Ancient Connections in Eurasia. Tübingen: Kerns Verlag. pp. 157-192. doi:10.51315/9783935751377.008 [citations: 5]
Books
2021 Reyes-Centeno H and Harvati K (Editors). Ancient Connections in Eurasia. Tübingen: Kerns Verlag. doi:10.51315/9783935751377
2019 Sahle Y, Reyes-Centeno H, and Bentz C (Editors). Modern Human Origins and Dispersal. Tübingen: Kerns Verlag.
2018 Harvati K, Jäger G, and Reyes-Centeno H (Editors). New Perspectives on the Peopling of the Americas. Tübingen: Kerns Verlag.
Reports - Published
2020 Rathmann H and Reyes-Centeno H. Das menschliche Gebiss als Spiegel unserer Evolution. Blickpunkt Archäologie 2:146-149.
Reports - Unpublished
2025 Keppeler J, Almeda FA, Bauzon L, Peterson J, Reyes-Centeno H. Report of archaeological fieldwork at the Kandingan Cave Limestone Area (Brgy. Amoslog, Placer, Surigao del Norte, Philippines). Submitted to the Museum of Anthropology, Archaeology Division, Manila: National Museum of the Philippines.
2024 Keppeler J, Almeda FA, Bauzon L, Peterson J, Reyes-Centeno H. Report of archaeological exploratory activities (July-August 2023). Submitted to the Museum of Anthropology, Archaeology Division, Manila: National Museum of the Philippines.
Presentations and Abstracts
2026 Benítez D, Delgado M, and Reyes-Centeno H. Patrones de parentesco y residencia posmarital de poblaciones Herrera y Muisca bajo un lente de genética cuantitativa y análisis de redes sociales. 2-5 June, Congreso Colombiano de Arqueología (CCA), Manizales, Colombia. [Podium]
2026 Brooks M and Reyes-Centeno H. Webb-Funkhouser: Legacy of anthropological thought at University of Kentucky. University of Kentucky Undergraduate Research Showcase. 29 April, Lexington, USA. [Poster]
2026 Gilmore L and Reyes-Centeno H. The First Rhode Island Regiment of 1777. University of Kentucky Undergraduate Research Showcase. 29 April, Lexington, USA. [Poster]
2026 Koontz S, Wise A, Nolin R, Keppeler J, Rodriguez M, Mink P, Crothers G, Murphy P, Robertson J, Uhl A, Bailey S, and Reyes-Centeno H. Integrating magnetometry and ground penetrating radar at the American Revolutionary War fortification in Butts Hill Fort (Portsmouth, Rhode Island, USA). International Conference on Computer Applications & Quantitative Methods in Archaeology (CAA). 31 March – 3 April 2026, University of Vienna, Austria. [Poster]
2026 Keppeler J, Göldner D, Copas C, Copas S, Peyroteo Stjerna R, Larena M, and Reyes-Centeno H. Inked in 3D: Surface modelling and image enhancement of a tattooed human mummified individual from Benguet, Philippines. International Conference on Computer Applications & Quantitative Methods in Archaeology (CAA). 31 March – 3 April 2026, University of Vienna, Austria. [Poster]
2025 Phillips KM and Reyes-Centeno H. Exploring the Denisovans and their impact on human evolution. Annual Meeting of the Kentucky Academy of Sciences. 21-22 November 2025, University of Louisville, USA. [Poster]
2025 Reyes-Centeno H. Mode and timing of human occupation in the Philippines. International Research Conference on Language and Community Heritage. 29-30 July, Palawan State University, Puerto Princesa, Philippines. [Podium]
2025 Reyes-Centeno H. Mode and timing of human occupation in Southeast Asia and Oceania. Oceanic and Southeast Asian Navigators (OCSEAN) 2025 International Conference. 22 July, Universitas Udayana, Denpasar, Indonesia. [Podium]
2025 Benítez D, Delgado M, and Reyes-Centeno H. The application of nonmetric dental traits in analyzing sex mobility and social networks among pre-Hispanic societies from the Northern Andes. 94th Annual Meeting of the American Association of Biological Anthropologists. 12-15 March, Baltimore, USA. American Journal of Biological Anthropology 186(S79):14-15, doi:10.1002/ajpa.70031. [Poster]
2025 Keppeler J and Reyes-Centeno H. UKinSPIRE: Community and collaborative approaches in the archaeology of the southern Philippines. Global Fair, University of Kentucky International Center. 12 February, Lexington, USA. [Poster]
2024 Keppeler J, Rodriguez M, and Murphy P, Reyes-Centeno H, Uhl A. Archaeological survey at Butts Hill Fort (Portsmouth, Rhode Island). 91st Annual Meeting of the Eastern States Archaeological Federation (ESAF). 7-10 November, Salve Regina University, Newport, Rhode Island, USA. [Podium]
2024 Rathmann H, Perretti S, Porcu V, Hanihara T, Scott RG, Irish J, Reyes-Centeno H, Ghirotto S, and Harvati K. Evaluating the utility of craniodental metric and non-metric data in biodistance analysis. Annual Meeting of the European Association of Archaeologists (EAA). 28-31 August, Sapienza University, Rome, Italy. [Podium]
2024 Crothers G, Keppeler J, Benítez D, and Reyes-Centeno H. 3-Dimensional Scanning of Human Trackways in Helms Deep Cave. National Speleological Society Convention. 5 July, Sewanee, Tennessee, USA. [Podium]
2024 Adel MM, Reyes-Centeno H, Beeman CS, Sharab L, Hartsfield JK. Precision assessment of facial asymmetry using 3D imaging and artificial intelligence. Annual Session of the American Association of Orthodontics (AAO). 3-6 May, New Orleans, USA. [Poster]
2024 Benítez D, Delgado M, and Reyes-Centeno H. Parentesco, patrones de residencia posmarital, y diversidad dental entre cacicazgos Muiscas del Norte de los Andes. Symposium: “Nueva Esperanza: Permanencias y convergencias del pasado en el presente,” 2-4 May, Congreso Colombiano de Arqueología (CCA), Santa Marta, Colombia. [Podium]
2024 Castro J and Reyes-Centeno H. A biocultural exploration of diet-driven mandibular variation over time. University of Kentucky Undergraduate Research Showcase. 24 April, Lexington, USA. [Poster]
2024 Keppeler J and Reyes-Centeno H. Classrooms to caves: Community and collaborative approaches in the archaeology of the southern Philippines. UNITE Research Showcase, University of Kentucky. 18 April, Lexington, USA. [Poster]
2024 Price S and Reyes-Centeno H. Dental morphological affinities of Denisovans, Neanderthals, and modern humans. University of Kentucky Undergraduate Research Showcase. 24 April, Lexington, USA. [Poster]
2024 Adel MM, Sharab L, Hartsfield JK, Beeman CS, Reyes-Centeno H. Precision assessment of facial asymmetry using 3D imaging and artificial intelligence. 51st Annual Moyers Symposium. 2-3 March, Ann Arbor, University of Michigan, USA. [Poster]
2024 Morford AL, Huffman RK, Sebastian L, Jacob J, Reyes-Centeno H, Ngan L, Hartsfield JK. Geometric morphometric analysis of non-White populations diagnosed with class-III-malocclusion. 2024 International Association for Dental, Oral, and Craniofacial Research / American Association for Dental, Oral, and Craniofacial Research / Canadian Association for Dental Research (IADR/AADOCR/CADR) General Session. 13-16 March, New Orleans, USA. [Poster]
2023 Reyes-Centeno H. The Negrito hypothesis and prospect for Mamanwa archaeology. II International Mamanwa Dialogue Symposium. 26 October, Surigao, Philippines. [Podium]
2023 Reyes-Centeno H. Bridging the past and present: Harnessing virtual archaeology for heritage conservation. OCSEAN Philippines International Conference on Language, Genes, Culture, History and Community Heritage II: Sustaining Heritage for Future Generations. 6-10 June, Tboli, Philippines. [Podium]
2023 Reyes-Centeno H. Mindanao at the crossroads: Toward resolving the Austronesian problem. OCSEAN Philippines International Conference on Language, Genes, Culture, History and Community Heritage I. 4-6 June, Davao City, Philippines. [Podium]
2023 Reyes-Centeno H, Baker CE, Balk J, Smith S, Seales B. EduceLab: Infrastructure for next-generation heritage science. Mid-scale Experimental Research Infrastructure Forum (MERIF) 2023. 22-24 May, Boston University, Boston, USA. [Poster]
2023 Twine SR, DeLisle I, Ingraham B, Lescault E, Rodriguez M, Uhl A, and Reyes-Centeno H. Analyzation of spring box artifacts in Newport, Rhode Island. Undergraduate Research Showcase, University of Kentucky. 26 April, Lexington, USA. [Poster]
2023 Athié Teruel B, Correia MA, Uhl A, Strauss A, and Reyes-Centeno H. Quantifying biological sex in ancient Brazil. Undergraduate Research Showcase, University of Kentucky. 26 April, Lexington, USA. [Poster]
2023 Athié Teruel B, Correia MA, Uhl A, Strauss A, and Reyes-Centeno H. Quantifying sexual dimorphism of the bony labyrinth and biological sex distribution in Lagoa Santa, Brazil. 92nd Annual Meeting of the American Association of Biological Anthropologists. 19-22 April, Reno, USA. [Poster]
2023 Benítez D, Delgado M, and Reyes-Centeno H. Kinship, postmarital residence patterns, and dental diversity among Muisca chiefdoms in the Northern Andes. 92nd Annual Meeting of the American Association of Biological Anthropologists. 19-22 April, Reno, USA. [Podium]
2023 Plemons A and Reyes-Centeno H. Improving identification of nonwhite decedents in forensic investigations. UNITE Research Showcase, University of Kentucky. 18 April, Lexington, USA. [Poster]
2023 Benítez D, Delgado M, and Reyes-Centeno H. Kinship, postmarital residence patterns, and dental diversity among Muisca chiefdoms in the Northern Andes. UNITE Research Showcase, University of Kentucky. 18 April, Lexington, USA. [Poster]
2023 Steenken MB, Bonzani RM, Endonino JC, Detisch MJ, and Reyes-Centeno H. Morphological and chemical signatures of Chenopodium: Application of optical and electron microscopy to seeds from experimental and archaeological contexts. 88th Annual Meeting of the Society for American Archaeology. 29 March – 2 April, Portland, USA. [Podium]
2023 Reyes-Centeno H. Genomic and craniodental phenomics. Workshop of the OCSEAN Consortium: Archaeology of Oceania and the Pacific. Pühajärve, Estonia, hosted by Tartu University. 8 March. [Podium]
2023 Sesma E and Keppeler J. Digging close to home: An archaeological field school in the university's back yard. Revisiting Global Archaeologies: Society for Historical Archaeology 2023 Conference on Historical and Underwater Archaeology (SHA) , Lisbon, Portugal, 4-7 January. [Poster] doi: 10.5281/zenodo.19476216
2022 Athié Teruel B and Reyes-Centeno H. Quantifying biological sex in ancient Brazil. University of Kentucky Board of Trustees Retreat. 20 October 2022, University of Kentucky, Lexington, USA. [Poster]
2022 Seales B, Crothers G, Reyes-Centeno H, Baker C, Chapman C. EduceLab: Midscale infrastructure for heritage science – A case study. Frontiers in Archaeological Sciences 3: Rethinking the Paradigm. 7-9 October 2022, Cornell University, Ithaca, USA. [Poster]
2022 Rathmann H, Perretti S, Porcu V, Ghirotto S, Reyes-Centeno H, Harvati K. Inferring human neutral genetic variation from craniodental phenotypes. Annual Meeting of the European Society for the study of Human Evolution. 21-25 September 2022, University of Tübingen, Germany. [Podium]
2022 Blair L, Reyes-Centeno H, Uhl A. Ontogenetic variation in human bony labyrinth morphology. Showcase of Undergraduate Scholars. 26 April, University of Kentucky, Lexington, USA. [Poster]
2021 Beaudet A, Reyes-Centeno H, d’Errico F, Backwell L, Wadley L, Zipfel B, de la Peña P. A revision of the Border Cave cranium. Human Evolution: From Fossils to Ancient and Modern Genomes (Virtual Conference). 2-4 November, Virtual from Wellcome Genome Campus, United Kingdom. [Podium]
2021 Rathmann H and Reyes-Centeno H. The utility of dental nonmetric ASUDAS traits and trait combinations for inferring human population history. 90th Annual Meeting of the American Association of Physical Anthropologists (AAPA). 7-28 April (on-line). [Podium] American Journal of Physical Anthropology 174(S71):88. doi:10.1002/ajpa.24262
2021 Reyes-Centeno H and Hefner TJ. Evolution of cranial macromorphoscopic trait variation in modern humans. Annual Meeting of the Human Biology Association. 29 March – 2 April 2021, virtual. [Poster], American Journal of Human Biology 33(S1):37. doi:10.1002/ajhb.23593
2020 Mori T, Profico A, Reyes-Centeno H, and Harvati K. Virtual reconstruction and geometric morphometric analysis of the mid-Pleistocene hominin KNM-OG 45500 (Olorgesailie, Kenya). Annual Meeting of the European Society for the study of Human Evolution. 24-25 September, on-line at eshe.eu. [Podium]
2020 Rathmann H and Reyes-Centeno H. The utility of different dental traits and trait combinations for inferring human population history. Annual Meeting of the European Society for the study of Human Evolution. 24-25 September, on-line at eshe.eu. [Podium]
2020 Camarós E, Carnicero S, Cabazón AR, Reyes-Centeno H, Göldner D, Vallejo-Llano J, Pereda E, Bolado R, and Arias P. Lethal violence or violent rituals during the Iron Age? A cross-disciplinary bioarchaeological and taphonomic analysis of a Cantabrian skull. 26th Annual Meeting of the European Association of Archaeologists. 26-30 August, Budapest, Hungary, switched to on-line at www.e-a-a.org [Poster]
